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Notes -
The broad thrust of this article is arguing against a strawman. Nobody really disagrees that Russians might have said NATO was a threat. Anyone in the West can point that out freely and openly without fear of reproach. The issue is that NATO wasn't actually a threat in any plausible scenario in the way that Russians were describing it. Russians (or Putin specifically) typically alluded to NATO aggression either from a ground invasion or a nuclear first-strike, both of which were never in the cards given it would start World War 3 and mean a huge portion of the Earth's population from both sides being wiped out in an instant. Some Russians may have drank the propaganda koolaid and genuinely believed the West was willing to eliminate Russia in a geopolitical equivalent of a murder-suicide, but they were mostly relegated to the fringes.
What Russians/Putin were actually worried about was one of three things:
Western cultural and economic hegemony. NATO expansion doesn't really directly impact this, but NATO expansion serves as a barometer that the West is still triumphing over the former Soviet Union.
The West fomenting pro-democracy movements in Russia, similar to the Color Revolutions. Much of Russian society and Putin in particular have a deep antipathy for democracy, seeing it as not only a personal threat but as an invasive, enemy ideology and incorrectly blaming it for the turmoil of the Yeltsin years. Again, this doesn't really have anything directly to do with NATO expansion, but the fact that NATO is expanding at all means the West is robust enough to possibly try something like a pro-democracy coup in the future.
Loss of their sphere of influence. Many Russians still see their country as a Great Power, and the fact that NATO even has the possibility of being extended to Ukraine is deeply insulting.
So yes, many Russians say "NATO is a threat". But no, no reasonable Russian thinks NATO is a threat in a conventional sense since Russia still has the largest nuclear stockpile in the world. Instead, saying "NATO is a threat" is used as a dogwhistle to stoke generalized anti-Western sentiment or to appeal to delusions of grandeur, i.e. that Russia should reassemble the borders of the Soviet Union.
Is it simply a "delusion of grandeur" to respond to threats even if not directly martial and only against one's dignity, culture, and preferred system of governance? That seems valid to me. I don't know if characterizing it as "generalized anti-Western sentiment" is correct either because I feel like the West would likely respond in a similar way in a similar situation (that is, the behavior is anti-Western but the motives are more universal than just being anti-Western).
The "delusion of grandeur" is in regards to many Russians wanting the days of being a great power back, when in reality the Russia of today is too dysfunctional, corrupt, and kleptocratic to keep up with the likes of the USA or China.
People in the West may very well have done similar things if roles were reversed. That doesn't necessarily make it just, correct, or rational though.
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