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I believe that's a slur against trans women rather than a more general criticism.
There's another meaning. It means that you have a weak frame. You hear some stuff that makes you feel something and you cave to pressure instead of maintaining that frame. You change something fundamental about yourself to please other people. In the worst, stupidest case, to please your critics or enemies.
There's a constant temptation to do this if you're on the right but you're not rural or working class. Everyone around you is constantly pressuring you. John Cornyn loses his Senate primary by 27 points, breaths a sigh of relief, and starts countersignalling Trump. His wife is happy. His kids are happy. The women in his wife's book club are happy. "So glad you came around John. Sure we don't like the crime and taxes, but Trump is such a meany. Did you hear about that kid they shot in Austin last week?"
It's so much easier to troon than to hold out against the pressure. Just means you have no frame.
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Using it as a general synonym of "go insane" is unorthodox but not totally unreasonable.
Oh, I use it to refer to when a leftist or theater kid gets inappropriately, expressively euphoric in a social setting.
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